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SISTEMA INTEGRAL DE INFORMACIÓN ACADÉMICA - PÚBLICO
Título del libro: Tree Of Life: Evolution And Classification Of Living Organisms Título del capítulo: Annelids Segmented Worms
Annelida, or segmented worms, is a morphologically diverse group of
animals, with a large number of species (around 22,000) that inhabit
practically all marine habitats, as well as many terrestrial and
freshwater environments. The placement of annelids in the tree of life
is still unclear, although most recent hypotheses relate them to
molluscs, nemerteans, phoronids, and brachiopods. The phylogeny within
this group also presents major challenges. Annelida has traditionally
been split into three classes: ``Polychaeta'' (bristleworms),
``Oligochaeta'' (earthworms), and Hirudinea (leeches); however, recent
phylogenetic analyses have shown that Hirudinea is nested within a
paraphyletic ``Oligochaeta'' forming a clade now referred to as
Clitellata. Furthermore, Clitellata also appears to be nested within a
paraphyletic ``Polychaeta.'' Additionally, other groups displaying a
morphology far from the typical annelid and considered as separate phyla
in the past, such as Echiura (spoon worms), Sipuncula (peanut worms),
Siboglinidae (pogonophores and vestimentiferans), and Myzostomida are
now also considered to be annelids, though there have been some
controversies about their specific position. This chapter offers an
updated synthesis of the evolution of segmented worms and particular
characteristics that were key to their evolution since the Cambrian.